Category: New Research

Walking with Dinosaurs (2025) featuring Ichnoplanet students

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Dinosaur documentaries have been a great way to communicate the science of paleontology and dinosaurs to a wide audience. Recently, Walking with Dinosaurs has returned with a new version since its original inception in 1999. This time, two Ichnoplanet students make an appearance in one of the episodes!

Jack Milligan and Kaitlin Lindblad feature as part of the field crew during the fifth episode of the new series, titled “The Journey North.” The episode centres on a megaherd of the ceratopsian Pachyrhinosaurus lakustai, as they migrate across the forests, floodplains and rivers of paleonorthern northwestern Alberta during the Campanian stage of the late Cretaceous period. On their journey, they face many dangers, such as infighting among their own, fearsome predators such as the tyrannosaur Gorgosaurus, and sudden flash floods.

Pictures from the Pipestone Creek Bonebed during filming of Walking with Dinosaurs. From left to right: an overhead view of a dense area of Pachyrhinosaurus bones, including limb bones, ribs, and skull material; A grid map created by the field crew prior to collecting the bones; a shot of two of the field crew being filmed by the camera crew. Photos by Jack Milligan, 2023

Jack and Kailtin were invited by Dr. Emily Bamforth, curator of the Philip J. Currie Dinosaur Museum in Wembley, Alberta, to partake in filming of the series, where they excavated bones of Pachyrhinosaurus from the famous Pipestone Creek Bonebed, which has produced thousands of bones since the 1970’s. Fossils that were found during the filming were small juvenile bones, a rib with a pathology, and large frill bones. All these bones were mapped and observed using both traditional and modern methods, including a grid square and 3D scanning software, respectively.

A segment of the episode shows Dr. Bamforth going to a tyrannosaur track site to take silicone moulds of the tracks to study their morphology and discuss the possible tracemaker. This site was published in PLOS One, “The dinosaur tracks of Tyrants Aisle: An Upper Cretaceous ichnofauna from Unit 4 of the Wapiti Formation (upper Campanian), Alberta, Canada” by Enriquez et al., 2022.

Walking with Dinosaurs is available now to buy on physical media and on streaming. Check out the official trailer and a behind-the-scenes featurette featuring Jack!

Listen to CBC Saskatoon Morning and CBC Saskatchewan The Morning Edition radio interviews with both Jack and Kaitlin talking about their experience being involved in the production of the show.

Written by Jack Milligan

Pichipoda manganae, a new armadillo trackway from the Neogene of Argentina.

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Former Ichnoplanet students Martín Farina, Verónica Krapovickas and co-authors have published a study on mammalian trackways from the Neogene Vinchina (Middle to Late Miocene) and Toro Negro (Late Miocene to Early Pleistocene) formations, in the La Rioja Province Vinchina Basin, Argentina!

Northwestern Argentina has an outstanding ichnological record of the Cenozoic vertebrate faunas. This study reports five mammalian trackways and the diversity of the trackmakers responsible. Of the four ichnospecies identified, three are new. The trackways come from deposits interpreted as floodplains of both meandering and anastomosed fluvial systems.

Pichipoda is a novel ichnogenus that has a didactyl to monodactyl manus and tridactyl mesaxonic pes impressions with robust digits with blunt tips. Two ichnospecies are assigned to Pichipoda, including P. manganae and P. vizcainoi. P. manganae (P. manganaei) is the largest ichnospecies of Pichipoda and has a didactyl manus, and the manus of P. vizcainoi shows an almost monodactyl morphology. Tridactylonicha marsicanae is another novel ichnogenera and ichnospecies, described by a tridactyl to didactyl paraxonic manus and tridactyl mesaxonic pes impression with long, slender, and pointed tips of the toe impressions. Dolichotichnus marae has tetradactyl paraxonic manus and tridactyl mesaxonic pes impressions.

Pichipoda manganae. Figure 2 from De La Fuente et al., 2025.

P. manganae and P. vizcainoi are interpreted as being produced by armadillos, with P. vizcainoi being attributed most likely to tolypeutines. Before this study, ichnofossils attributable to fossil armadillos were unknown, meaning this is the beginning of our understanding of the trace fossil record of this group and can help us to ask further paleobiological questions. T. marsicanae is interpreted as being produced by hegetotheriids, a family belonging to the extinct group of South American ungulates, the Notoungulata. D. marae is most likely produced by dolichotines, a group of caviid rodents.

Astute readers will notice the etymology of P. manganae honours our very own Dr. Gabriela Mángano, which we here at Ichnoplanet firmly endorse. The contributions Dr. Mángano has made to ichnology are unmatched, and it is fitting that these newly described trace fossils act as a reflection and reminder of her great career.

Congratulations to the authors, including Martin and Verónica, on this amazing study. We know the continued research into the small mammal trackways of Cenozoic Argentina is filling in an important gap in our understanding of vertebrate faunas within South America, and we await what new discoveries have to say.

Read the paper in the Journal of South American Earth Sciences here.

Written by Jack Milligan

Congrats to Dr. Brittany Laing!

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We would love to congratulate Dr. Brittany Laing on her recent successful PhD defence! Her thesis focused on the behavioural evolution and ecological impact of organisms as seen through trace fossils, specifically applying theories and methodologies from mathematics & behavioural biology to trace fossil datasets in order to quantitatively examine behavioural and ecological changes through time. She also completed a master’s degree with our research group where she studied the ichnology at the Ediacaran and Cambrian GSSP in Newfoundland, Canada. She is also to thank for the Ichnoplanet website!

We are forever grateful for your contributions and wish you all the best in your future endeavours!

Written by Jack Milligan

Meet Himatiichnus mangano!

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Recently, an international team of ichnologists led by Katherine A. Turk described a new ichnogenus from the late Ediacaran Huns Member of the Urusis Formation in southern Namibia. They described a series of meandering, intertwining tubes exhibiting dual lineation patterns preserved in convex relief, appearing to disappear into the sediment and reemerge on the sediment surface. The distal terminal ends of the tubes are rounded and bulbous. Modern priapulid worms have produced such structures via peristaltic probing. This is similar to the famous Ediacaran-Cambrian biostratigraphic marker trace fossil Treptichnus pedum, also attributed to priapulid worms.

Figure 2 from Turk et al., 2024 showing the holotype of Himatiichnus mangano. See paper here.

The team also noted that these new trace fossils represented the earliest occurrence of Scalidophora, and are much like those traces of worms in the Cambrian, suggesting ecological interactions typical of the Cambrian Explosion appeared earlier than previously thought (the long fuse hypothesis). The morphological evidence pointed to these trace fossils being a novel ichnogenus and ichnospecies. The team gave the trace fossils the name Himatiichnus mangano, named after our very own Dr. M. Gabriela Mángano!

We couldn’t agree more with this decision, as Dr. Mángano has not only spent her career furthering the field of ichnology from all aspects, but has also contributed greatly to research on the Ediacaran-Cambrian transition around the world. Congrats, Dr. Mangano, on this great honour and achievement!

Get the scientific paper in Royal Society Open Science here.

Written by Jack Milligan

Grasslands National Park Fieldwork 2024

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Towards the end of the summer, I had the opportunity to continue fieldwork at two multitaxic Edmontosaurus quarries within the Uppermost Cretaceous Frenchman Formation in the East Block of Grasslands National Park in southern Saskatchewan under the supervision of Dr. Emily Bamforth of the Philip J. Currie Dinosaur Museum and Dr. Ryan McKellar of the Royal Saskatchewan Museum. These excavations have been ongoing and have yielded some fantastic fossil specimens and interesting geology.

We collected a limb bone (tibia) from one of our two Edmontosaurus quarries, called the “Leo” quarry. The tibia shows feeding traces as evidenced by well-defined tooth marks. During the excavation of the tibia, a shed tooth from a tyrannosaurid (Tyrannosaurus rex) was collected from the same depositional layer. This association of tooth traces on Edmontosaurus bones and T. rex teeth suggests that Leo was scavenged during subaerial exposure on a floodplain within a meandering fluvial system. Limb bones have a high likelihood of being consumed due to large locomotor muscles being present (a great food source for predators like T. rex), and thus will tend to have more tooth traces on the bone.

On an adjacent butte within the same depositional layer as the “Leo” quarry, there is a point bar deposit containing a second active quarry named the “Clifford“ quarry. While the Edmontosaurus in this quarry has yet to produce elements with tooth marks, teeth from T. rex and the crocodyliform Borealosuchus have shown up this past season. Adding to the mystery of this quarry, several articulated turtle skeletons have been collected from this deposit, showing no evidence of being scavenged.

Tooth marks have been a widely debated aspect of vertebrate ichnology. Recently, the category-modifier (CM) system was proposed by Wyenberg-Henzler et al., 2024, as a method to identify different morphologies of tooth marks and their associated behaviours. While not strictly defined within a traditional ichnological framework, it is the most comprehensive overview of tooth marks on bones in the literature to date. Based on the CM system, the tooth marks left on the Edmontosaurus tibia appear to be internally chattered, referring to the crescentic depression on the inside of the tooth mark. Shed carnivore teeth from both quarries suggest that the dominant predators within this ecosystem were crocodyliforms and tyrannosaurs.

Further excavation of the quarries is required, and research into the taphonomy of these two Edmontosaurus quarries is ongoing, but there are sure to be some exciting discoveries to be made in future field seasons!

Written by Jack Milligan